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The latter is a competitor of 2-Oxoglutarate-dependent dioxygenases such as Ten-eleven translocation methylcytosine dioxygenase and therefore induces DNA hydroxymethylation and gene upregulation 52

Similarly, glutathione has been reported to repress sulfur assimilation and uptake activities of sulfate transporters in a number of species (Buchner et al., 2004)